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I click on a random link (it is a valid URL) and it returns a 404. It doesn't matter where the link comes from, put it in the browser address bar and get a 404. Refresh the page again and again over a period of time still a 404.
In another browser tab go to the website main URL and then drill down through the directory structure and find the page in question. It is there and valid, no error.
Both URLs are identical.
If I now go back into the browser tab with the 404 page and refresh I now get the correct page displayed!