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characters Google will treat as seperators in the URL

     
8:44 pm on May 11, 2005 (gmt 0)

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Despite the debates that perennial reoccur is seems pretty clear to me that Google treats dashes (-) in URL's as seperators and underscores (_) as part of the word.

My question is are there other characters that Google treats like seperators maybe tilde (~), bang(!), or comma (,).

How well do these work with other search engines?