I'm hoping to start a new thread solely focused on Penguin recovery tips
I think it's premature to speculate about how best to recover, when we know almost nothing about Penguin.
Here are a few of the things we don't know:
Whether Penguin removes the benefit of certain tactics or "unnatural" patterns, or whether Penguin penalizes sites having those "unnatural" patterns. (The difference is important for how one tries to recover from Penguin.)
Whether the tactics in question are reflected in the site itself (e.g. internal links) or external to the site (reflected in the inbound link profile).
Whether the tactics in question only become problematic when they are combined with other tactics.
Whether the tactics in question only become problematic when they are used in excess (too much of a good thing).
Despite the length of the other thread, there was no consensus on even the most basic issues -- and surprisingly little speculation, or specific clues that might lead to any sort of meaningful list of recovery tips.